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Do the 200A and 400A Crouse-Hinds Arktite plugs and receptacles, allow larger wire well size connectors to interchange with connectors of other wire well sizes?

Yes the larger wire well sizes for the 200A and 400A Arktite Plugs and Receptacles will mate with the smaller wire well sizes as long as the grounding style 1 or style 2 and the wires/poles on each are identical.